Ficus ixoroides is a rare rheophytic fig only found along rivers in Sarawak and northern Kalimantan. The fig fruits ripen green/yellow and look similar to Ficus fistulosa which suggests that the ripe female figs attract bats for seed dispersal.
Secondary dispersal: However the thick narrow leaves and rheophytic riverine habit suggests that female fig fruit containing seeds are also dispersed by water and fish. Rivers always flow downhill so all river or stream dispersed plants also require a secondary disperser (in this case bats and possibly fish) which can carry the seeds upstream and uphill.
Photos by Astrid Cruaud and Jean-Yves Rasplus.Location: River bank between Pa Lungan to Pa Umor in the Bario Highlands of NE Sarawak. Co-ordinates: 3.761009 115.526397