PHOTO ABOVE: This Ficus virens growing on the floodplain of the Tempassuk river at Kota Belud in Sabah is believed to have the widest canopy (70 m) of any tree in Borneo.
INTRODUCTION TO THE 6 SECTION UROSTIGMA STRANGLERS FOUND IN BORNEO
World 27 spp. Borneo: 6 spp.
Borneo hosts 6 species of Section Urostigma stranglers including 2 of Borneo’s biggest fig trees F. caulocarpa and F. virens and the introduced Bo Tree (F. religiosa) sacred to Buddhists, Hindus and Jains. None are endemic. All species are monoecious (bisexual). For recent taxonomy see;
Chanturasuwan (2013) A revision of Section Urostigma
The 2 most common Section Urostigma stranglers in Borneo Ficus virens and Ficus caulocarpa are easily distinguished from each other. Ficus caulocarpa always has reddish brown petioles (leaf stalks) whilst with Ficus virens the petioles are always light green.
SPECIES | STATUS | RIPENS |
F. caulocarpa | Common especially coastal districts | White spotted pink |
F. concinna | One record Bod Gaya island, Semporna | White spotted purple |
F. prasinicarpa | Local on Derawan islands Kalimantan | Green to purple |
F. religiosa | Introduced Hindu/ Buddhist temples | Reddish green |
F. superba | South China Sea islands only | Purple spotted white |
F. virens | Common throughout | White spotted purple |
How to distinguish Section Urostigma stranglers from Section Conosycea stranglers.
- Fig fruits normally with short stalks ( F. religiosa which has no stalk (peduncle) is an exception)
- 4 species of Section Conosycea fig fruits also have stalks but these are all bat figs which ripen green.
- Urostigma fig fruits ripen white first later developing pink or purple spots. Again F. religiosa is an exception as the figs ripen reddish green) Conosycea figs tend to ripen yellow/orange/red/black without spotting. ( F. microcarpa ripens white pink purple black but is not spotted)
- Fig fruits are normally ramiflorus, densely packed on old growth branches. Conosycea figs normally grow in the leaf axils
- Articulated leaves with long stalks (petioles) with hinges or elbows (pulvinus) at both ends enabling the leaf to flex in the wind.
- Inside Section Urostigma figs the male stamens are clustered around the ostiole in all Borneo species whilst with Section Conosycea stranglers the male stamens are spread throughout the interior of the fig fruit.
- All are deciduous, shedding their leaves in one event which are then immediately replaced.
- The twigs show evidence of seasonal intermittent growth by changes in the length and color of the internodes.
- The intermittent seasonal growth is reflected in differences in the stipules of Sections Conosycea and Section Urostigma. Dormant Urostigma stipules are short blunt,conical and brown.
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Dormant (resting) stipule of Ficus caulocarpa photographed by Shuai LIAO at Sukau, Kinabatangan River in Sabah. Collection #2019477.